Mutation-generated variation in behavior is thought to promote the evolution of cooperation. Here, we study this by distinguishing two effects of mutation in evolutionary games of the finitely repeated Prisoner’s Dilemma in infinite asexual populations. First, we show how cooperation can...

There is continuing debate about what explains cooperation and self-sacrifice in nature and in particular in humans. This paper suggests a new way to think about this famous problem. I argue that, for an evolutionary biologist as well as a quantitative social scientist, the triangle of two...

This paper addresses the question of whether our evolutionary history suggests that humans are likely to be individually selected selfish maximizers or group selected altruists. It surveys models from the literature of evolutionary biology in which groups are formed and dissolved and where the...

In a haystack-type representation of a heterogeneous population that is evolving according to a payoff structure of a prisoner’s dilemma game, migration is modeled as a process of “swapping” individuals between heterogeneous groups of constant size after a random allocation fills the...