Most of the literature on the evolution of human pro-sociality looks at reasons why evolution made us not play the Nash equilibrium in prisoners' dilemmas or public goods games. We suggest that in order to understand human morality, and human prosocial behaviour, we should look at reasons why...

The main ingredient of this paper is the derivation of the generalized version of Hamilton's rule. This version is derived with the Generalized Price equation. The generalized version of Hamilton's rule generalizes the original rule, in the sense that it produces a set of rules; one rule for...

The main ingredient of this paper is the derivation of the Generalized Price equation. This generalizes the original Price equation in the sense that it produces a set of Price-like equations, one for every different underlying model that one could assume has generated the data. All of these...

Repeated games tend to have large sets of equilibria. We also know that in the repeated prisoners dilemma there is a profusion of neutrally stable strategies, but no strategy that is evolutionarily stable. This paper shows that for all of these neutrally stable strategies there is a stepping...

A widespread claim in evolutionary theory is that every group selection model can be recast in terms of inclusive fitness. Although there are interesting classes of group selection models for which this is possible, we show that it is not true in general. With a simple set of group selection...